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In a world abandoned by God, the bodies of the dead will continue to walk unless they are buried by one of the mysterious Gravekeepers. Ai, one of the last children in a world where no more can be born, becomes her small village's Gravekeeper when her mother, the previous Gravekeeper, passes on… but the arrival of a pale gun-wielding stranger destroys her village and what little remained of Ai's old existence. Is this seemingly immortal man possibly an important figure from Ai's past?
Uncertain of what else to do, Ai finds herself compelled to set forth on a mission to learn the secrets of this strange world without proper death, unravel the mysteries of her own past, and, hopefully, find a way to put the dead permanently to rest!