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With skills honed during World War II when they worked undercover at the site of the United Kingdom's main decryption establishment, the women of Bletchley Circle use their code-breaking skills to solve crimes in post-war London. Joining Anna Maxwell Martin (Susan), Rachael Stirling (Millie), Sophie Rundle (Lucy), and Julie Graham (Jean), who are all returning from Season 1, is Olivier Award-nominated actress Hattie Morahan (Alice).
Two mysteries are featured this season. In "Blood on Their Hands," former Bletchley Park girl Alice Merren is in prison awaiting trial for the murder of eminent scientist, John Richards. Jean stands behind her innocence and tries to reunite the circle in an attempt to help one of their own. Lucy is now getting on well in a clerical job at Scotland Yard and keeping her head down, and Millie is working as a German translator. Susan is reluctant to get involved after her experience with the twisted serial killer Malcolm Crowley and the danger it posed to her family.
In "Uncustomed Goods," Millie branches out into what she believes to be a harmless job on the side dealing in uncustomed goods. This endeavor leads to her being kidnapped and held in a seedy hotel by some gangsters. On discovering this, Alice goes to Jean and Lucy to help. The girls start to investigate in order to save their friend and, as they do so, uncover a much darker, hidden world of crime.